Dr.Dodson 9.25.96 Autonomic nervous system Practical aspects. made of sympathetic and parasympathetic components. there are two convenient concepts to recall. all organs of the body have both sympathetic and parasympathetic innervation. the nerves tend to go to the organs as a mixed nerve, with both symp and parasymp fibers in it. These tend to be small unmyelinated nerves. Sympathetic and parasympathetic differ greatly, though. Parasympathetic = craniosacral system Sympathetic = thoracolumbar system this separation is interesting - the sympathetic system arises in the thoracid and lumber region, coming off the spinal cord in each thoracic and lumbar segment, in successive segments. You can't destroy the sympathetic system, therefore, because there are many segments. the parasympathetic arises in the head, distributing as cranial nerves, and the sacral region, distributing as the pelvic nerve. So, the sympathetic system needs to get to the head, and the parasymp needs to get to the body. how? travel together in the neck as the vagus nerve (cranial nerve X) vagus distributes parasympathetic fibers to heart stomach and gut. There is a "switchboard" called the cranial-cervical ganglion which separates the vagus and sympathetic systems at the base of the head. Also, middle cervical ganglion splits them at the thoracic inlet. vagus splits off of trunk, and splits into dorsal and ventral, travelling through diaphragm. sympathetic splits into greater/lesser splanchnic which form a plexus (cranial mesenteric plexus, where the fibers of the vagus nerve join, so the fibers leaving the plexus are mixed sympathetic/parasympathetic nerves, which then distribute to the organs. note that after the splanchnic nerves come out, the sympathetic trunk decreases in size, but it does continue, through to the caudal part of the animal. another splanchnic (lumbar splanchnic) nerve splits off in the lumbar region, and forms another ganglion called the caudal mesenteric ganglion. after this split the sympathetic trunk dies out, but sympathetic fibers are carried into the pelvis via the hypogastric nerve, which comes out from the ganglion and travels into the pelvis. there is yet another plexus (pelvic plexus) in region of vagina/prostate. The pelvic and hypogastric nerves run into this plexus, mixing symp and parasymp fibers, and then these mixed nerves innervate bladder, rectum, etc. So, all pelvic viscera recieve this mixed nerve supply. the pelvic nerve is dorsal in position, not very conspicuous, hard to find in dissection. Easiest thing to do is find caudal mesenteric plexus (near caudal mes. artery) and watch hypogastric nerve running caudally from it into the pelvis. Key pelvic artery= vaginal or prostatic artery. pelvic plexus should be caudal to that artery. If you find the pelvic plexus, you might find the pelvic nerve dorsally, leading into the pelvic plexus. it's not one of the great wonders of nature or anything, so don't stress out over it. Illustration in small millers fig 105 pg 136, fig 143 p 199, fig 150 p 212. dr.dodson recommends coloring in the pix.